The terrestrial ecosystem model. In this figure GPP, RA, LC, and RH represent gross primary production, autotrophic respiration, litterfall, and heterotrophic respiration, respectively. These carbon fluxes are in grams of carbon per square meter per month (g C m−2 month−1). CV, CS, NVL, NVS, NS, and NAV represent mean annual values of vegetation carbon, soil carbon, labile nitrogen within vegetation, structural nitrogen within vegetation, soil nitrogen, and inorganic nitrogen in soils, respectively. The mean annual carbon values are in grams of carbon per square meter (g C m−2), while nitrogen values are in grams of nitrogen per square meter (g N m−2). NRESORB, NMOBIL, LN, NUPTAKEL, NUPTAKES, NETNMIN, NINPUT, and NLOST represent the amounts of resorbed nitrogen as labile nitrogen from structural nitrogen, nitrogen mobilized for production via recycling, nitrogen in litter production, nitrogen uptake by vegetation for labile nitrogen, nitrogen uptake by vegetation for structural nitrogen, net nitrogen mineralization, nitrogen input to the ecosystem, and nitrogen lost from the ecosystem, respectively. These nitrogen fluxes are in grams of nitrogen per square meter per month (g N m−2 month−1).